Set at the dawning of World War 2, humans have begun to disappear. Some come back, some do not. Madelyn Bryce is one such person who is chosen to take the train to an undisclosed location, but her upcoming death is not the sort she was expecting.

While humankind is at the brink of turmoil, the Vampyres have their own troubles--it is rumored that the detestable species known as Lycans has been one-upped by a new problem breed: Hybrids that are both Vampyres and Lycans. Not all of the Vampyres agree that such abominations should be done away with and they seek to keep them safe from the disapproving Councils.